Entry 1 sense Webster, 1913 Metathorax /mĕtˌ-əth-ôr'-ăksˌ/ · Met·a·tho·rax · IPA /mˌɛtəθˈɔrˌæks/ 01 n. The last or posterior segment of the thorax in insects. See Illust. of Coleoptera. 1. The last or posterior segment of the thorax in insects. See Illust. of Coleoptera.(Zool.) See: Coleoptera